The film is inspired by the fable The Sky is Falling (also known as Chicken Licken, Henny Penny, or Chicken Little). In the small suburban town of Oakey Oaks, Chicken Little rings the school bell and cries for everyone to "run for your lives!" This sends the whole town into a frenzied panic that causes so much havoc that it destroys a small part of the town; eventually they calm down enough to ask him what's wrong, and Chicken Little explains that a piece of the sky shaped like a stop sign had fallen on his head when he was sitting under the big oak tree in the town square. He's unable to find the piece now. His father, Buck Cluck, ashamedly assumes that this "piece of sky" was just an acorn that had fallen off the tree and had hit him on the head. Chicken Little becomes the laughing stock of the town.
A year later, Little has become famous in the town for being crazy, which has led to people avoiding him because of his supposed insanity, his only friends are outcasts like himself: Abby Mallard, who is called "Ugly Duckling" (who has a crush on Chicken Little); Runt of the Litter, who is extremely large despite being the smallest in his family; and Fish out of Water, who wears a helmet full of water and does not speak. All have been mocked and teased by neighborhood bully Foxy Loxy and her friend Goosey Loosey.
During a dodgeball game (the coach pits the popular vs. the unpopular), Abby tells Little to talk to his dad, when Little wants his dad to be proud. While Little, Abby and Fish are dodging, Runt is getting hit by every ball thrown at him. To help Chicken, Abby hands Little magazines about talking to your parents, but he hands them to Fish, who immediately starts ripping pages out of each one, making a miniature building and airplanes. Fish then proceeds to pull a King Kong impersonation. Foxy and Goosey bully Little and Friends soon after.
Chicken Little joins his school's Little League baseball team in an attempt to recover his reputation and his father's pride, but is unfairly made last, while Foxy Loxy impresses spectators and the newspapers with her pitches and "miracle catches" -- until the ninth inning when Chicken Little, reluctantly called to bat by the coach, who tells him not to swing as he's certain that if Chicken Little tries to swing, he'll lose the game for them, scores an inside-the-park home run after two strikes.
But that night back at home, while celebrating his victory by singing "We Are The Champions" by Queen in his own style, and his father's praise, he is hit on the head - by what appears to be a chunk of the sky shaped like a stop sign - only (after narrowly avoiding telling his father about it) to find out that it is not a piece of the sky, but something else. It has chameleonic characteristics- it's not invisible, but it blends into the background (which would thereby explain why Chicken Little was unable to find it last time). Meanwhile his friends are singing karaoke to the Spice Girls' "Wannabe", until Little calls them over to help figure out what it is.
When Fish pushes a button on the back of the octagon, it flies back up into the sky, taking Fish with it. It turns out to be part of the camouflage of an invisible alien spacecraft. They chase the flying ship until it lands; they go inside, encounter a small orange alien, find Fish, and are about to escape when two aliens in robotic suits spot them. A chase ensues that goes outside the ship and into town (the orange alien still following in secret). Little manages to ring the bell to warn everyone, but the aliens see the crowds coming and manage to escape, leaving the orange one behind. No one believes the story of an alien invasion; Little is ridiculed all over again, and everything seems to be going down the tubes for him, until the next day; he and his friends discover the orange alien, and a few minutes later a whole fleet of alien ships descends on the town.
The invasion is actually a misunderstanding, as two aliens (who are red and yellow making their child orange) are looking for their lost child (whose name is Kirby) and attack only out of concern. It's not an invasion, it's a rescue mission. As the aliens rampage throughout Oakey Oaks, vapourising people and objects (among them Foxy Loxy and the mayor) seemingly at random, Little realizes he must return Kirby to his parents to save the planet. At first he tries to do it covertly but once the aliens attack, he's forced to confront his father and regain his trust first. They share a caring moment in a movie theater. Abby interrupts them and requests that they hurry up. Little and Cluck are seen leaving the theater, but before leaving, Chicken Little admits his romantic feelings for Abby and kisses her.
In the invasion, Buck Cluck, now regaining his pride and trust in Little, defends Little from the aliens, even going as far as deflecting an incoming vaporiser ray with a garbage can lid, throwing the lid at the creatures, and punching them away at the top of town hall. Abby, Runt and Fish also aid Chicken Little in his quest to return Kirby to his parents. It is then discovered that the aliens weren't vaporising people, the ray guns teleported them aboard a spaceship. Afterwards, the aliens return everything to normal (though Foxy Loxy's mind was slightly scrambled after she was restored, making her more girly and kind), and Hollywood makes a heavily dramatized film about Chicken Little. The actor portraying Little is a handsome, muscular rooster while Abby is an attractive looking goose. Runt is a brutish hog with horns who dies a heroic death while fighting the aliens and the actor who plays Fish looks exactly the same in the same but is able to converse in English. At the end of the movie, Little and Abby are seen secretly holding hands in their popcorn bucket, and everyone cheers for Chicken Little's efforts to save the town.
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